Tanjirou Kamado dan teman-temannya Pembasmi Iblis menemani Hashira Api Kyoujirou Rengoku, untuk menyelidiki kasus misterius yang terjadi di dalam kereta. Tak banyak yang mereka ketahui tentang keberadaan Enmu, salah satu dari Dua Belas Kizuki yang juga berada di dalam kereta dan telah menyiapkan perangkap untuk mereka.
It is the Taisho Period in Japan. Tanjiro, a kindhearted boy who sells charcoal for a living, finds his family slaughtered by a demon. To make matters worse, his younger sister Nezuko, the sole survivor, has been transformed into a demon herself. Though devastated by this grim reality, Tanjiro resolves to become a “demon slayer” so that he can turn his sister back into a human, and kill the demon that massacred his family.
The curtain rises on Sarasa Watanabe, a starry-eyed 5'10" student who hopes to perform as a male lead role in the all-female Kouka Acting Troupe. She forms a friendship with her new roommate, a former idol. Together, they’re in for the role of a lifetime.
